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Virology is an area of study within the Division of Medical Sciences, an administrative unit based at Harvard Medical School that coordinates biomedical PhD activities at the Longwood Medical Area. Students who study in Virology receive a PhD in medical sciences. Prospective students apply through Harvard Griffin GSAS; in the online application, select “Division of Medical Sciences” as your program choice and select "Virology" in the area of study menu.

Virology is one of the programs in the Harvard Integrated Life Sciences that facilitates collaboration and cross-disciplinary research. Visit HILS for additional  application instructions .

This program is one of the few virology graduate programs in the country, and its small size provides the benefits of a smaller program, offering nearly a one-to-one ratio of students to faculty.

You will take advantage of a well-developed curriculum that focuses on analyzing, reading, and discussing papers to generate, present, and discuss research proposals. Most importantly, you will learn how to think as a scientist. You will have the opportunity to participate in state-of-the-art research involving molecular biology, cell biology of viruses, structural analysis, cryo-EM, and genomic analysis of cells and viruses.

Examples of student projects include structure and mechanism actions of antiviral antibodies including SARS-CoV2, cellular genes that promote and inhibit viral infection, and mechanisms of viral oncogenesis.

Graduates have gone on to faculty positions at prestigious institutions such as MIT, Brandeis University, Duke University, and Yale University. Others have begun careers at leading companies like Moderna, Gilead, Pfizer, and Sanofi Pasteur.

Personal Statement

Standardized tests.

GRE General: Not Accepted GRE Subject: Not Accepted iBT TOEFL minimum score: 100 IELTS minimum score: 7

See list of Virology faculty

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Virology and Gene Therapy

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Virology and Gene Therapy Track

faculty-to-student ratio

Bench-to-bedside thesis topics spanning basic virology and translational virology

Guaranteed 5-year internal fellowship.

includes full tuition, stipend and benefits

As outbreaks of potentially deadly diseases like influenza, Ebola or West Nile Virus continue to make headlines, so does the need to study the pathogens that cause them. Virologists play a key role in biological science, working to dissect and understand the nature of viruses and translate that knowledge into clinical practice. Discoveries over the past few decades show great promise in areas such as vaccine development, gene therapy and immunotherapy.

The Virology and Gene Therapy Track within the Ph.D. Program at Mayo Clinic Graduate School of Biomedical Science offers a highly productive, interactive research environment for you to develop as an independent investigator. As a student, you’ll learn from and work alongside faculty members who have primary interests in virology, viral vectors and gene therapy. These areas overlap with the fields of biochemistry, cell and molecular biology, genetics, and immunology.

Current areas of research include:

• Molecular biology of viruses
• Mechanisms of virus-host interactions
• Gene therapy
• Oncolytic virotherapy
• Cancer immunotherapy
• Vaccine development
• Tissue engineering using viruses
• Genetic engineering using viruses

Students receive a comprehensive education in the biomedical sciences through a set of core courses. Specialized tutorials and journal clubs provide advanced training in the broad areas of molecular virology, host-cell interactions, tumor immunology, gene therapy of metabolic diseases, cancer gene therapy and vector development.

Students are introduced to the laboratories participating in the program. You have the opportunity to visit these laboratories and select three in which you spend eight weeks participating in a research project. You'll select your thesis lab in the spring.

In conjunction with the laboratory rotations, you begin fulfilling the core curriculum requirements as well as the virology and gene therapy requirements. Most students complete the core courses by the end of their first year, in addition to taking the written qualifying exam.

As a second-year student, most of your time is spent in the lab developing preliminary data toward your thesis project. By December of the second year, you draft your thesis proposal and take the oral qualifying examination on your proposal.

Second-year students also take advanced tutorials in virology and gene therapy as well as related areas.

The third and subsequent years are devoted primarily to pursuing thesis research with some additional courses.

Together with a thesis adviser, you select faculty members to participate in your thesis advisory committee. Thesis committee meetings assess the trajectory and evaluate the progress of your thesis research project on a regular basis. Upon completion, you write a thesis and present your findings in seminar form. This is followed by a thesis defense.

The Virology and Gene Therapy Track is a one-of-a-kind program that spans basic research in viral vectors to downstream analysis of clinical trial samples. This track prepares you to gain a solid understanding of virology as well as preclinical and clinical product development. Students in this track have a unique opportunity to see firsthand how academia interfaces with clinical, biotechnical and industrial interests to bring the next therapeutics from concept to patient bedside.

Justin Maroun M.D.-Ph.D. student, Virology and Gene Therapy Track

Mayo Clinic offered me the unique opportunity to study the biology of viruses and how to genetically alter them to become gene therapy vectors for my graduate studies. Mayo is probably is the only institution in the country that offers a graduate program this specialized.

Jeffrey Rubin Ph.D. student, Virology and Gene Therapy Track

I chose the Virology and Gene Therapy Track based on my interest in molecular virology and infectious disease. Our faculty not only has expertise in molecular virology, but we have experts in the field of oncolytic virotherapy and gene therapy as well. We also have access to patient samples, collaborations across the institution, and phenomenal core facilities.

Crystal Mendoza Ph.D. student, Virology and Gene Therapy Track

I was a tech at Mayo before transitioning to graduate school. I witnessed firsthand the value of having research buildings located alongside clinic buildings. Collaboration exists not only across departments but also within the clinic. We have clinicians attend our lab meetings, and I have clinicians on my thesis committee to help guide my research into actual treatments.

Christopher Driscoll Ph.D. student, Virology and Gene Therapy Track

Recent thesis topics

• “Use of Glucokinase Gene Delivery to Enhance Beta-Cell Proliferation and Function,” Brian Lu, Ph.D. (Mentor: Yasuhiro Ikeda, Ph.D.)
• “Sensing of HIV-1 by the Innate Immune System,” Swati Kumar, Ph.D. (Mentor: David Dingli, M.D., Ph.D.)
• “The Innate Immune System is a Major Determinant for Successful Oncolytic Measles Virotherapy," Cheyne B. Kurokawa, Ph.D. (Mentor: Evanthia Galanis, M.D.)
• "The Dual Role of Perforin in the Balance Between Protection and Pathology During CNS Viral Infection and Blood-Brain Barrier (BBB) Disruption," Robin C. Willenbring, Ph.D. (Mentor: Aaron Johnson, Ph.D.)
• “B-type Natriuretic Peptide: Biology and Therapeutic Applications," Sara J Holditch, Ph.D. (Mentor: Yasuhiro Ikeda, Ph.D.)
• “Evaluation of Viral Gene Expression and E3 Immunomodulatory Functions of Adenovirus Serotype 26 to Inform Vector Design for Cancer Therapy," Mallory A. M. Turner, Ph.D. (Mentor: Michael Barry, Ph.D.)
• "Characterizing and Advancing Oncolytic Measles Virus Therapy Against Lymphoma," Tanner S. Miest, M.D., Ph.D. (Mentor: Roberto Cattaneo, Ph.D.)
• "Engineering and Development of Single Cycle Adenovirus Vectors as Mucosal Vaccination Platforms," Catherine M. Crosby, Ph.D. (Mentor: Michael Barry, Ph.D.)

Your future

Many graduates of the Virology and Gene Therapy Track choose to pursue postdoctoral training regardless of whether they intend to pursue careers in academia or industry. Other students choose to enter advanced training programs like clinical microbiology and biochemical genetics programs.

After graduating from the program, you could also choose to pursue a career in education, scientific writing and editing, or become a scientific grant program officer. Several students from our laboratories have become tenured faculty and leaders in industry and in foundations.

Meet the director

Welcome to the Virology and Gene Therapy track at Mayo Clinic — a leading medical institution where you’ll receive training from some of the world’s brightest, most-distinguished scientists and physicians.

Our program works with other research and clinical programs at Mayo to facilitate rapid bench-to-bedside translation as well as easy access to clinical samples.

Our mission is to provide high-quality education you won’t find anywhere else.

Michael Barry, Ph.D. Virology and Gene Therapy Track Director Professor of Medicine Phone: 507-266-9090 Email: [email protected] See research interests

Browse a list of Virology and Gene Therapy Track faculty members

Immunology, Microbiology and Virology PhD Program

The application for fall 2024 is now open.

The Immunology, Microbiology, and Virology (IMV) Program is designed for students interested in obtaining a Ph.D. in Microbiology & Immunology. 

We offer in-depth coursework and diverse research opportunities that focus on immunology, microbiology and virology.

The flexibility of our training program allows students to train in a number of exciting  research areas  including: cancer, autoimmune disease, respiratory pathogens and often allows students to develop highly effective interdisciplinary collaborations, resulting in cutting edge thesis projects.

Housed in the  Department of Microbiology and Immunology , the Immunology, Microbiology, and Virology PhD Program (IMV) houses a diverse group of faculty with expertise in bacteriology, virology, fungi, and immunology. Students interested in pursuing a doctoral degree in these disciplines should  apply  to the program.

Learn more about our program Student Perspectives Alumni Perspectives

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What Sets Us Apart

Learn more about the core courses, electives, and lab rotations that will be part of the IMV curriculum.

We support and guide trainees in several areas of research. Browse the current research our faculty and trainees are doing by research area.

Mentor Relationships

Our students most commonly reference the personal relationships and valuable mentoring they receive as one of the top reasons why they recommend URMC.

Discover Microbiology & Immunology at URMC

Latest News

June 20, 2024 What Is H5N1 Bird Flu, How it Spreads to Humans, and Safety Concerns

September 19, 2023 A Game Plan for Timing Your Flu, Covid and RSV Shots This Fall with Commentary by David Topham

April 17, 2023 Wilmot Researchers Discover How to Steer Army of Immune Cells toward Cancer

Upcoming Events

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  Thu, Sep 05 @ 12:00 PM   MC | K307 (3-6408)

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What Our Students and Alumni Say...

"The mentorship and the exceptional research experiences I had in the IMV program prepared me for my academic career. The program really does a great job of addressing many of the skills you will need to be successful once you complete your training. Aspects of the program that have been invaluable to my career were: the opportunity to present my work to an audience regularly, the exceptional classes (that I still rely on to teach my own coursework!), the responsibility of developing and completing a research project of the highest caliber and the support of the faculty in providing expert insight into my work and its potential. As a student I was held at a very high standard. I would not have been able to set up my own lab (at Rollins), make the right decisions regarding my research and the management of my lab if it weren’t for my training at UR."

Brendaliz Santiago-Narvaez, PhD

Dr. Santiago-Narvaez is currently an Assistant Professor in the Department of Biology at Rollins College in Winter Park, Florida. She teaches courses for the Biology and Biochemistry/ Molecular Biology Majors and also has her own independent research lab.

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About the PhD in Molecular Microbiology and Immunology Program

Through a departmental core curriculum as well as research area-specific courses, laboratory rotations, qualifying examinations, mentored research, and professional development, all MMI PhD students are prepared to engage in cutting edge research and scholarship that advances knowledge. MMI PhD students hone their scholarship, research and professional skills. Course and laboratory work can address problems in microbial pathogenesis, immunology, disease transmission, and diseases related to malaria, mosquito and arboviral biology.

Students can choose to complete the traditional MMI PhD program or the MMI PhD program concentration in Rigorous, Reproducible, and Responsible Research Investigation in Immunology & Microbiology (R 3 IM). The concentration in Rigorous, Reproducible, and Responsible Research Investigation in Immunology & Microbiology (R 3 IM) conveys a broad background in immunology and infectious diseases research, with a special emphasis on critical thinking, logic, ethics, and written and oral skills to help graduates become communicators of complex scientific concepts and agents of change in their workplaces and communities.

What Can You Do With a Graduate Degree In Molecular Microbiology And Immunology?

Visit the  Graduate Employment Outcomes Dashboard to learn about Bloomberg School graduates' employment status, sector, and salaries.

Sample Careers

• Postdoctoral Fellow, Research Associate
• Science Communicator or Writer
• Scientist (academia, industry)
• Specialist in Science Regulatory Affairs
• Faculty, Professor
• Science Advocate (nonprofit agencies)
• Public Health Service (NIH, CDC, FDA)
• Science Policy Fellow

Curriculum for the PhD in Molecular Microbiology and Immunology

Browse an overview of the requirements for this PhD program in the JHU  Academic Catalogue  and explore all course offerings in the Bloomberg School  Course Directory .

Current students can view the MMI student handbook on the MMI portal site.

Topic Areas

MMI faculty are recognized experts in a wide variety of infectious diseases research areas, allowing our PhD students to study the biology of disease based on their research interests and career goals. Our PhD students gain a comprehensive understanding of infectious diseases that provides a foundation to launch careers that directly tackle critical public health challenges.  

Topic Areas include: 

• Bacterial pathogenesis
• Cell biology 
• Fungal pathogenesis 
• Malaria 
• Medical entomology/Disease ecology
• Parasite pathogenesis 
• Rigor, Reproducibility, and Responsibility in Scientific Practice
• Tick-borne diseases
• Vaccine development 
• Vector biology
• Viral pathogenesis

Admissions Requirements

For general admissions requirements, please visit the How to Apply page.

Standardized Test Scores

Standardized test scores (GRE) are  optional  for this program. The admissions committee will make no assumptions if a standardized test score is omitted from an application, but will require evidence of quantitative/analytical ability through other application components such as academic transcripts and/or supplemental questions.  Applications will be reviewed holistically based on all application components.

Vivien Thomas PhD Scholars

The  Vivien Thomas Scholars Initiative (VTSI)  is an endowed fellowship program at Johns Hopkins for PhD students in STEM fields. It provides full tuition, stipend, and benefits while also providing targeted mentoring, networking, community, and professional development opportunities. Students who have attended a historically Black college and university (HBCU) or other minority serving institution (MSI) for undergraduate study are eligible to apply. To be considered for the VTSI, you will need to submit a SOPHAS application, VTSI supplementary materials, and all supporting documents (letters, transcripts, and test scores) by December 1, 2024. VTSI applicants are eligible for an  application fee waiver , but the fee waiver must be requested by November 15, 2024 and prior to submission of the SOPHAS application.

Per the Collective Bargaining Agreement (CBA) with the JHU PhD Union, the minimum guaranteed 2025-2026 academic year stipend is $50,000 for all PhD students with a 4% increase the following year. Tuition, fees, and medical benefits are provided, including health insurance premiums for PhD student’s children and spouses of international students, depending on visa type. The minimum stipend and tuition coverage is guaranteed for at least the first four years of a BSPH PhD program; specific amounts and the number of years supported, as well as work expectations related to that stipend will vary across departments and funding source. Please refer to the  CBA to review specific benefits, compensation, and other terms

Need-Based Relocation Grants Students who  are admitted to PhD programs at JHU starting in Fall 2023 or beyond can apply to receive a need-based grant to offset the costs of relocating to be able to attend JHU.   These grants provide funding to a portion of incoming students who, without this money, may otherwise not be able to afford to relocate to JHU for their PhD program. This is not a merit-based grant. Applications will be evaluated solely based on financial need.  View more information about the need-based relocation grants for PhD students .

Questions about the program? We're happy to help. Ashley Choi  Senior Academic Coordinator Alex Kim  Senior Academic Program Coordinator  [email protected]

Microbiology and Molecular Genetics Program at Emory University

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About the Program

Emory University's Graduate Program in Microbiology and Molecular Genetics trains students to use microorganisms as models for investigating fundamental problems in genetics.

  This interdisciplinary program offers training in a variety of areas, from microbiology and molecular biology to cell biology and genetics. The curriculum emphasizes the application of modern approaches to fundamental problems in microbial genetics and molecular biology.

Throughout this program, students will have the chance to engage in rigorous training in state-of-the-art laboratory procedures and participate in courses on bacterial genetics and physiology, microbial development, molecular biology of viruses and bacterial pathogens—as well as mechanisms of pathogenesis against bacteria and viral diseases.

The program is intended for students interested in medicine and will prepare them for careers working with pharmaceutical companies or other medical-related organizations.

Bacteriology

Offers research opportunities that  provides a multidisciplinary environment for studies in bacteriology with a central focus on molecular mechanisms of pathogenesis and antibiotic resistance. Specific areas of research include  bioinformatic approaches to bacterial virulence,  regulation of gene expression, mechanisms of intracellular survival, and much more.

Offers research opportunities that explore the  various areas of virology, with particular strengths in research on HIV, influenza, and the large DNA viruses. Covers a broad range of basic research topics such as virus entry, replication and assembly, aspects of virus-host relationships such as mechanisms of pathogenesis and immune responses of the host, and more translational areas of focus involving drug design and vaccine development.

Parasitology

Offers research opportunities that  focus on malaria host-pathogen interactions, with the aim of better understanding how malaria infection takes hold and causes disease. Research topics include mechanisms of parasite invasion and residence in host cells, antigenic variation, immune mechanisms of parasite control, immunopathogenesis of malaria infection and the impact of co-infections on malaria.

MMG graduates are highly sought after by employers in various industries, including pharmaceutical companies, biotechnology firms, and government agencies. They work as researchers, laboratory managers, and technical specialists in the industry and pursue independent research careers at universities or medical schools.

Life at Emory & Atlanta

Life at Emory University is one of the most exciting and intellectually stimulating experiences you could ask for.

A diverse, inclusive community welcomes students and faculty from all over the world. You'll find plenty of opportunities to get involved in campus life—and many resources available that will help you thrive as an Emory student.

The university has consistently ranked among the top 20 institutions nationally for NIH research support and was named one of the "New Ivies" by Newsweek—a tribute to both its academic excellence, as well as dedication to teaching.

Our faculty members are some of the most distinguished scholars in their fields and work closely with undergraduate students on research projects, independent study courses, or senior thesis projects.

Located just 6 miles northeast of downtown Atlanta, Emory's 704-acre campus is in a beautifully wooded area that offers students a peaceful environment to study but also provides easy access to Atlanta's bustling cultural scene.

Inside the Program

Latest News Stories

Graduate Program

Virology Ph.D. degree will give students the opportunity to participate in state-of-the-art research involving molecular biology, cell biology of viruses, structural analysis, cryo-EM, and genomic analysis of cells and viruses.

• Current Students

Infectious Diseases and Immunity PhD

The IDI is a laboratory-based research program where students study infectious diseases and immunology through a public health lens.

The IDI Program intends to create opportunities for students to gain new and advanced knowledge about infectious disease agents and how they interact with host cells, human populations, and the environment. Our goal is to improve public health by increasing our understanding of infectious diseases and human immunology through basic and translational research that contributes to developing new diagnostics, treatments, and methods to prevent or control diseases. Because the IDI Graduate Group is administratively managed by the School of Public Health, we follow the School of Public Health application process and deadlines . GRE test scores are not required for the Infectious Diseases and Immunology PhD program.

Program Objectives

The objective of this program is to provide students with research-oriented training that will enable them to design and implement independent investigations and advance the fundamental knowledge of infectious disease agents and their interactions with the human host and the environment. The IDI PhD program is a five to six-year program. Students in the program are fully funded throughout their time in the program, and are provided with a competitive monthly stipend.

We strive to promote health by integrating basic research and applied technologies to develop new approaches for the diagnosis, treatment, prevention, and control of infectious disease in humans. This program combines clinical, epidemiological, and basic laboratory research strategies in order to apply these methods to specific infectious disease problems affecting human populations. In addition, students have opportunities to interact with faculty members from multidisciplinary centers involved in global infectious disease research. These include the Center for Global Public Health and the Center for Emerging and Neglected Diseases. Students matriculating through this program will acquire expertise in not only fundamental infectious disease research, but also learn how their research relates to other disciplines. Students matriculating through this degree program will acquire expertise in fundamental infectious diseases research for which there is demand from academic institutions, local and national government agencies, and biotechnology companies.

Our Commitment to Belonging

The graduate students and faculty of the Infectious Disease and Immunity Program stand with Black Lives Matter. We believe that racism and police violence is a public health crisis that deserves the same amount of attention and work as any other disease.

As students of Public Health and Biology, we understand the ways in which our field and institutions have been complicit and contributed to the harm perpetuated against Black bodies. The popularized pseudoscience of phrenology, the grotesque distortion of evolution into eugenics, and the irreparable harm of the Tuskegee Syphilis study are just a few examples of how science and public health have perpetuated white supremacy in both mind and body.

We are dedicated to creating an academic environment free of anti-Blackness and making our program and community places where Black, Indigenous, and other URM students can thrive. This is a lofty goal and we recognize that it will take more than a passing attempt to root out anti-Blackness from the academy and ourselves. We pledge to make this effort a sustained one, creating accountability by updating publicly the progress we have made on our goals and setting new goals year after year.

• DEI Resources on campus Includes campus initiatives, affinity groups, STEM outreach in Bay Area
• Grants/fellowships and funding A consolidated list of of funding and fellowship opportunities

Program Snapshot

• First year: Coursework and three lab rotations
• End of first year: Join dissertation research lab
• Second year: Complete coursework requirement, GSI for a semester and take qualifying examination (QE)
• Third year: Advance to candidacy after passing the QE
• 3rd-5th/6th years: Hold dissertation committee meetings at least once a year, complete the annual Doctoral Progress review, complete dissertation research, complete GSI one additional time. Present dissertation at IDI Monday Seminar. Notify program their graduating semester. File dissertation.

IDI students spend their first year taking an interdisciplinary set of classes and completing three lab rotations. The curriculum aims to introduce students to the breadth of infectious disease and immunology research while sharpening statistical skills and building a strong foundation of basic science knowledge. These courses are supplemented by a small faculty led seminar in the first and second years that focuses on developing and improving specific skills like grant writing, qualifying exam preparation and critical research paper analyses.

For additional details on program requirements, see curriculum information and resources for current students page.

Qualifications

The following subjects are normally required as undergraduate preparation for all candidates. Deficiencies must be made up early during the graduate program.

• Mathematics: Calculus; one course in probability or statistics.
• Physics: General physics.
• Chemistry and biochemistry: Inorganic chemistry; organic chemistry; biochemistry; and associated laboratories.
• Biology: General biology lecture and laboratory; genetics; and a basic course(s) in molecular biology.
• Common undergraduate majors for admitted applicants: Biology, integrative biology, microbiology, biological sciences, biology and communications
• Common work experience for admitted applicants: Work experience is not required for admission, but relevant work experience related to infectious diseases, e.g. wet laboratory and/or surveillance work is important for the IDI PhD program. Most of the admitted applicants have strong lab and research experience.

GRE scores are not required, it is optional for the fall 2023 admissions cycle. We recommend submitting a GRE if you have no other evidence of quantitative, verbal, or analytical abilities in your application.

Recruitment Information

The Infectious Diseases and Immunity PhD program admits students only in the Fall semester. Applications are available for submission online in mid-September. The deadline for applications is December 4, 2023, for Fall 2024 admission. Late applications are not accepted. Students should complete their applications as early as possible and be sure to hit the submit button before the deadline. Admission review will be conducted in mid January 2024. Shortlisted applicants will be invited to attend the required interview on Feb 9, 2024.

IDI Graduate Group faculty come from multiple departments including Infectious Diseases and Vaccinology, Plant and Microbial Biology, and Molecular and Cell Biology. In addition, a unique aspect of the IDI program is our affiliation with UCSF faculty who conduct work in the area of global infectious disease. IDI students may choose to join any IDI affiliated lab for their thesis research.

UCSF Affiliated Faculty

Below is a list of UCSF professors who are currently affiliated with IDI but who are only a small selection of potential UCSF mentors.

Margaret Feeney, MD

Bryan Greenhouse, MD, MA

Phil Rosenthal, MD

Rachel Rutishauser, MD, PhD

Current IDI PhD Students

Student Name – Faculty Advisor; Lab

• Cuong Joseph Tran – Dr. Matthew Welch ; The Welch Lab
• Joanna Vinden – Dr. Bryan Greenhouse ; Greenhouse Lab (UCSF)
• Marcus Wong – Dr. Eva Harris ; The Harris Research Program
• Eric Jedel – Dr. Suzanne Fleiszig ; Fleiszig – Evans Lab
• Kishen Patel – Dr. Kim Seed ; Seed Lab
• Reinaldo Mercado-Hernandez – Dr. Eva Harris ; The Harris Research Program
• Elias Michael Duarte – Dr. Eva Harris ; The Harris Research Program
• Abigail Kane – Dr. Rachel Rutishauser ; Rutishauser Lab, UCSF
• Claire Mastrangelo – Dr. Lee Riley ; Riley Lab
• Carolina Agudelo – Dr. Ashley R. Wolf, co-mentor Dr. Sarah Stanley ; Stanley Lab
• Isabel Lamb-Echegaray – Dr. Sarah Stanley ; Stanley Lab
• Jaime Cardona Ospina – Dr. Eva Harris ; The Harris Research Program
• Scott Espich
• Felix Pahmeier – Dr. Eva Harris ; The Harris Research Program
• Marize Rizkalla
• Zahra Zubair-Nizami

Go to “Curriculum Information and Resources for Current IDI Students” page

We would like to thank the following funding donors for their generous fellowship support to support our students and our work:

• Hillel and Rose Levine Fellowship
• The Albert and Mildred Krueger Memorial Fellowship
• Sally Anne Bradley Presser and Steven A. Presser Fellowships

Prospective donors: please visit give.berkeley.edu .

Biomedical Graduate Studies

Cell & molecular biology graduate group.

Stephen Bart, Bates lab

• Microbiology, Virology, and Parasitology

Welcome to the Microbiology, Virology, and Parasitology program!

Infectious diseases resulting from viruses, parasites, prions, and bacteria are a major cause of human morbidity and mortality. Some important infectious diseases, including HIV, malaria, and hepatitis C are becoming more rather than less prevalent. The threat of emerging infectious diseases and bioterrorism also calls for increased research in the area of microbiology, and in fact the NIH is greatly increasing research funding for work on infectious diseases. The recent outbreak of SARS and the continued spread of West Nile virus in North America are but two recent examples of emerging infectious diseases. By studying human pathogens, it is also frequently possible to learn much about normal cell biology, molecular biology, and immunology - infectious agents have long been used as model systems to study important processes.

The Microbiology, Virology, and Parasitology program provides students an opportunity to undertake concentrated study in the molecular and cellular biology of viral and bacterial pathogenesis and parasitology. Program faculty conduct research in a broad range of disciplines, including Bacteriology, Emerging Infectious Diseases, Immune Response, Microbial Genomics and Evolution, Parasitology, Tumor Virology, Virology, and Host Microbiome. Viruses, parasites, prions, and bacteria are a major cause of human morbidity and mortality. Disease resulting from HIV, malaria, and tuberculosis are becoming more prevalent. The threat of emerging infectious diseases, such as Zika and West Nile viruses, and bioterrorism also calls for increased research in the area of microbiology. Students study human pathogens, as well as their interplay with host resident microbial populations, learning much about normal cell biology, molecular biology, and immunology, as well as developing strategies for the prevention and treatment of infectious diseases.

The program has an extensive series of seminars designed to not only expose students to the latest and hottest research in microbiology, but to give students an opportunity to present their work to a large and diverse audience. The program feels that it is important for students to gain experience in speaking about their work in public, as this is an important facet of any job in science, and a weekly Tuesday noon seminar with average attendance of approximately 90 faculty, students, postdocs and technicians provides this forum for the virology trainees. Similar seminar series are held for our bacteriology and parasitology students. There's also a Presentation and Communication Review (PCR) monthly series for all MVP students, organized by MVP students, to further strengthen and practice science communication. The Wednesday Microbiology seminar series features prominent scientists from throughout the country and Europe who talk about their latest work in virology, bacteriology, parasitology, and immune responses. As part of this series each semester, there is an Alumni Day when a former MVP program student or postdoc who is now an Assistant Professor at another institution returns to campus to talk about their work and to meet with current students over lunch to talk about their careers. More information on various seminars can be found on the Seminar Series page .

Molecular Virology

About the Program

Study the viral life cycle or the comorbidities initiated by viral infection when you pursue your PhD in Molecular Virology at Case Western Reserve, one of the top research universities in the U.S. Our interdisciplinary approach to biomedical sciences, through our Biomedical Sciences Training Program , allows you to focus on molecular virology while also taking electives in related disciplines. With that background, you’ll have a strong foundation to pursue a career as a scientist in the biotech field, work as a technical specialist, or continue on to a postdoctoral fellowship. Plus, Cleveland is a great place to advance your career—there are more than 700 biotechnology and biomedical companies in town as well as four top medical institutions.

Student Resources

Whether you’re looking for information about education abroad opportunities, have questions about visas, or are interested in international opportunities on campus, these quicklinks will help you quickly navigate some of the key resources our website offers for students.

What to Expect

Start your coursework in cell and molecular biology and advance from there to virology and the host response to viral infections. Smaller class sizes mean you’ll receive individualized attention and mentorship during your six years (or fewer) here.

Then put your knowledge to the test in the lab, where you’ll conduct your research. You can gain funding for HIV/AIDS research in our Center For Aids Research, which is supported through a T32 institutional research training grant. Beyond Ohio, we have an international research collaboration with colleagues in Kampala, Uganda.  

Request Information

Beyond the classroom.

As you pursue your research interests here, you’ll have opportunities to show off your work—locally and internationally. Whether you choose to go to the Gordon Conferences, the American Society for Microbiology or another convention, our faculty will be there to support you as you prepare your presentation. If you’re more interested in attending seminars rather than presenting, check out the Lester O. Krampitz Seminars and the Center For Aids Research’s Leaders in AIDS speaker program, both of which host top researchers.

You’ll also have dozens of on-campus clubs to choose from. The Bug Club, for example, hosts chats on virology, microbiology, antibiotic resistance and parasites, while other School of Medicine Journal Club participants prompt discussions on HIV/AIDs, immunology and more.  

By the Numbers

tuition coverage, including healthcare

medical research school in the U.S., U.S. News & World Report

private institution in federal expenditures for research and development, National Science Foundation

in the world for contributions to innovation, Nature Index Innovation

faculty-to-student ratio

Admissions Requirements

Our program thrives when we have students with diverse perspectives and a range of educational, professional and life experiences. We look forward to learning more about you!

The application process requires:

• Online application , including a statement of purpose
• $50 application fee
• Bachelor’s degree and/or master’s degree from an accredited institution (either already complete, or will be complete before you enroll)
• Unofficial transcripts
• Current resume/CV
• List of research experiences (optional)
• Three letters of recommendation
• International applicants: Results of TOEFL, IELTS or PTE  

Quick Links

Degree Programs

The Department of Immunology and Infectious Diseases focuses on the biological, immunological, epidemiological, and ecological aspects of viral, bacterial, and protozoan diseases of animals and humans and the vectors that transmit some of these infectious agents. Research in the department emphasizes basic pathogenic mechanisms that may lead to better diagnostic tools, the development of vaccines and other interventions for prevention and control of infection and disease, and the identification of new targets for antiviral and antiparasitic drugs. Research in the department is described extensively on our “About the department” page and individual faculty pages.

The department provides a unique multidisciplinary training environment for PhD students. Students dive deeply into a specific aspect of immunology and infectious disease in their own thesis research while gaining broad exposure through interactions with our close community in the department and across the School. In a typical week, a student whose research focuses on the genetics of TB drug resistance might also attend a departmental work-in-progress talk from a student working on immune responses to HIV and a departmental seminar from an outside leader reporting progress on a TB control program from a specific country. Many students also have the opportunity to travel within the US and globally as part of collaborative research projects.

PhD students in our department join through PhD programs in the “Harvard Integrated Life Sciences” (HILS) consortium. We’ve included the most common programs below. You can see a full list of HILS programs here .

Ph.D. Programs

Ph.d. program in biological sciences in public health.

For more information about the PhD program visit the Division of Biological Sciences

PhD Program in Biological and Biomedical Sciences

For more information about the PhD program visit the The Biological and Biomedical Sciences (BBS) Program

Ph.D. Program in Immunology

Immunology and molecular biology of parasitic and other infections This area of interest introduces students to recent advances in the biology of parasitic and infectious diseases and provides background for conducting research on these diseases. The program emphasizes molecular biology, immunology, cell biology, and the epidemiology of parasites.

Infectious disease epidemiology and tropical public health This area of interest provides a solid understanding of epidemiology, ecology, and control of infectious diseases in developing countries. It emphasizes control and prevention measures and the biological basis of diseases caused by pathogens that range from viruses to parasites.

For more information about the PhD program visit the Ph.D. Program in Immunology

Ph.D. Program in Virology

For more information about the PhD program visit the Ph.D. Program in Virology

Vector Biology, Ecology, and Control

This area of interest focuses on the manner in which blood-feeding arthropods interact with their various vertebrate hosts and with the human pathogens that they transmit. These interests combine biological experimentation, epidemiological analysis, and population studies.

Students become familiar with the various arthropods that are associated with human disease and learn the ways environmental change may result in ill health. Students conduct studies on mechanisms of transmission of vectorborne pathogens, both in the laboratory and in the field, and devise novel intervention strategies.

MD-PhD programs

Our PhD students are also eligible to join supplemental concentrations that provide close communities with additional opportunities for scientific training, career and professional development, mentorship, and friendships. We’ve included a few popular programs below. You can see a full list here .

• Harvard Graduate Program in Bacteriology
• Harvard Infectious Diseases Consortium
• Harvard Therapeutics Graduate Program
• Leder Human Biology and Translational Medicine Program
• HSPH Interdisciplinary Concentrations (e.g. Infectious Disease Epidemiology)
• University-wide Secondary Fields (e.g. Science, Technology, and Society)

We encourage applicants from all backgrounds and identities. To learn more about application resources, fee waivers, affinity groups, and other supports for Harvard PhD students, please visit this page .

News from the School

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How for-profit medicine is harming health care

A tradition of mentoring

Promising HIV treatment

We have 31 Virology PhD Projects, Programmes & Scholarships

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Virology PhD Projects, Programmes & Scholarships

A PhD in Virology would provide you with the chance to lead your own research project that will further our current understanding of viruses. Whether you’re researching the response of the human immune system to a given virus, attempting to use viruses to deliver therapeutics, or working on a vaccine, you’ll be aiming to improve and save lives.

What’s it like to do a PhD in Virology? 

Doing a PhD in Virology, you’ll become proficient in the laboratory with techniques including fluorescent antibody staining, polymerase chain reaction (PCR) and cell culture. You’ll spend time reading around your research area and gain inspiration for methods to improve your experimental work.

Some typical research topics in Virology include:

• Investigating viral replication
• Assessing the impact of co-infection on an organism
• Vaccine development
• Developing novel therapies using viruses
• Improving methods of viral detection
• Studying the immune response to a certain virus

Almost all Virology PhD projects are advertised with the main research goal pre-determined. Many of these projects come with full funding attached, though some may require you to self-fund. Finding your own funding can be challenging since you must pay both PhD and bench fees. Self-funding and the difficulty finding a supervisor with research interest similar to yours, who also has the right equipment for your work.

During an ordinary workday, you’ll be in the laboratory performing, preparing or planning experiments, writing up sections of your thesis, and chatting to your colleagues and supervisor about your current and upcoming work.

To be awarded your PhD, you’ll need to submit a thesis of approximately 60,000 words and you’ll defend this during your viva exam .

Entry requirements

The  entry requirements  for most Virology PhD programmes involve a Masters in a subject directly related to Biology, with some experience in microbiology, at Merit or Distinction level. If English isn’t your first language, you’ll also need to show that you have the right level of  language proficiency . 

PhD in Virology funding options 

The research council responsible for funding Virology PhDs in the UK is the Biotechnology and Biological Sciences Research Council (BBSRC). They provide fully-funded studentships including a stipend for living costs, a consumables budget for bench fees and a tuition fee waiver. Students don’t apply directly to the BBSRC, you apply for advertised projects with this funding attached.

It’s uncommon for Virology PhD students to be ‘self-funded’ due to the additional bench fees. However, if you were planning to fund yourself it might be achievable (depending on your project) through the UK government’s PhD loan and part-time work.

PhD student (f/m/d) in Molecular Cell Biology/Virology in Göttingen and Bordeaux

Phd research project.

PhD Research Projects are advertised opportunities to examine a pre-defined topic or answer a stated research question. Some projects may also provide scope for you to propose your own ideas and approaches.

Funded PhD Project (Students Worldwide)

This project has funding attached, subject to eligibility criteria. Applications for the project are welcome from all suitably qualified candidates, but its funding may be restricted to a limited set of nationalities. You should check the project and department details for more information.

The role of neutrophils in the airway during bacteria- viral co-infection in COPD

Self-funded phd students only.

This project does not have funding attached. You will need to have your own means of paying fees and living costs and / or seek separate funding from student finance, charities or trusts.

FULLY FUNDED - The epidemiological underpinnings of viral spillover risk from bats

Funded phd project (uk students only).

This research project has funding attached. It is only available to UK citizens or those who have been resident in the UK for a period of 3 years or more. Some projects, which are funded by charities or by the universities themselves may have more stringent restrictions.

Predicting the Evolution of Influenza and SARS-CoV-2 Viruses

Selection and evaluation of new universal influenza vaccine candidates using monoclonal antibodies and digitally designed ha and na libraries & pseudotype virus screens, development of a dna amplification-free assay for pathogen detection, uts phd scholarship- sydney water- crcsaafe ( domestic only), exploring mechanisms underlying viral infections in initiating pulmonary fibrosis, competition funded phd project (students worldwide).

This project is in competition for funding with other projects. Usually the project which receives the best applicant will be successful. Unsuccessful projects may still go ahead as self-funded opportunities. Applications for the project are welcome from all suitably qualified candidates, but potential funding may be restricted to a limited set of nationalities. You should check the project and department details for more information.

Funded PhD- Physiology and therapeutic modulation of oxidative phosphorylation in HIV-1 infected cells.

The use of machine learning models to identify new non-nucleoside reverse transcriptase inhibitors targeting hiv-1, the development of new antiviral agents targeting hiv-1, control and inhibition of virus replication, self-funded phd- integrated multi-omics characterisation of human retroviral reservoirs, self-funded phd– role of t and nk-cell immunity during dengue virus infection and impact of immune modulation during therapy.

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• Perelman School of Medicine /

Cell and Molecular Biology: Microbiology, Virology, and Parasitology, PhD

Related programs.

• Cell and Molecular Biology: Cancer Biology, PhD
• Cell and Molecular Biology: Cell Biology, Physiology, and Metabolism, PhD
• Cell and Molecular Biology: Developmental, Stem Cell, and Regenerative Biology, PhD
• Cell and Molecular Biology: Gene Therapy and Vaccines, PhD
• Cell and Molecular Biology: Genetics and Epigenetics, PhD

Cell and Molecular Biology

The Cell and Molecular Biology Graduate Group (CAMB) is an interdisciplinary graduate program, providing rigorous training in modern cell and molecular biology, preparing students for leadership careers in biomedical research. Within this integrated program are six discipline areas:  Cancer Biology ;  Cell Biology, Physiology, and Metabolism ;  Developmental, Stem Cell and Regenerative Biology ;  Gene Therapy and Vaccines ;  Genetics and Epigenetics ; and  Microbiology, Virology and Parasitology . Program faculty include more than 300 scientists representing 35 departments from the Perelman School of Medicine, the Schools of Arts and Sciences, Dental Medicine, and Veterinary Medicine, Children’s Hospital of Philadelphia, the Wistar Institute and Fox Chase Cancer Center. The research efforts of these scientists are diverse in their focus, experimental system, methodology, and represent the leading edge of basic and translational biomedical science.

Students from colleges and universities around the nation and the world are enrolled in the program, selecting one discipline area based on their scientific interests, yet have access to the full breadth of curricular and research opportunities provided by this large and diverse program. Our students participate in core courses in cell and molecular biology, specialized coursework in one or more discipline areas, and original hypothesis-driven thesis research. Upon completion of the PhD, they pursue successful research careers at top academic institutions, in the biotech and pharmaceutical industries, and in other biomedicine-related career paths.

For more information:  http://www.med.upenn.edu/camb/

Microbiology, Virology, and Parasitology

The Program in Microbiology, Virology, and Parasitology provides students an opportunity to undertake concentrated study in the molecular and cellular biology of viral and bacterial pathogenesis and parasitology. Program faculty conduct research in a broad range of disciplines, including Bacteriology, Emerging Infectious Diseases, Immune Response, Microbial Genomics and Evolution, Parasitology, Tumor Virology, Virology, and Host Microbiome. Viruses, parasites, prions, and bacteria are a major cause of human morbidity and mortality. Disease resulting from HIV, malaria, and tuberculosis are becoming more prevalent. The threat of emerging infectious diseases, such as Zika and West Nile viruses, and bioterrorism also calls for increased research in the area of microbiology. Students study human pathogens, as well as their interplay with host resident microbial populations, learning much about normal cell biology, molecular biology, and immunology, as well as developing strategies for the prevention and treatment of infectious diseases.

For more information: https://www.med.upenn.edu/camb/mvp.shtml

View the University’s Academic Rules for PhD Programs .

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• PhD in Infectious Diseases

A competitive graduate program with an emphasis on existing and emerging interdisciplinary research areas

For those interested in a research-driven career in academia or industry, our Department of Infectious Diseases offers a PhD program with training in many disease relevant disciplines such as molecular virology, pathogenic bacteriology, classical and molecular parasitology, epidemiology, immunology, avian disease, and wildlife disease.

Important aspects of our graduate training program include:

• A seminar series that features excellent speakers from around the country
• Five journal clubs that cover the range of department research interests and provide important training in critical analysis of published work
• A yearly department retreat that offers an opportunity for our graduate students to present their work to their peers
• Competitive financial support

In addition to a range of molecular biology and immunology tools, the department has equipment to support phosphoimaging, quantitative real-time PCR, flow cytometry, computational bioinformatics, and chemiluminescent gel imaging.

Additionally, our college’s close proximity to the Centers for Disease Control, the U.S. Department of Agriculture, and the Environmental Protection Agency offices, as well as our access to facilities such as UGA’s Animal Health and Research Center, the Center for Vaccines and Immunology, and Diagnostic Laboratories, make us uniquely positioned for collaboration and innovation in education and research.

About the program

How it works.

Applicants are accepted through the  University of Georgia’s Integrated Life Sciences (ILS) program . The ILS program allows first-year graduate students to explore the research areas of more than 200+ faculty and 14 participating units before they choose a major professor and departmental home.

Eligibility

Competitive applicants will have:

• Bachelor’s degree or master’s degree in any biological science from an accredited institution (For international applicants, an equivalent to American bachelor’s degree requiring a minimum of four years of training is required)
• Demonstrated research experience

For a list of current tuition and fees, see  UGA’s Bursar & Treasury Services . Financial assistance is available from the Department of Infectious Diseases. A majority of our students are funded by research and teaching assistantships, training grants, and fellowships. Please contact the department directly for more information.

How to apply

Step 1: An online application must be completed through the  UGA Graduate School. This will include the submission of official college transcripts; a résumé or curriculum vita; three letters of recommendation; and a personal statement describing your experiences, interest in infectious diseases, and your reasons for applying to the program.

Foreign applicants are required to submit additional documentation as part of their online application. See the  Graduate School’s  website for a list of supplemental information for your country.

Step 2: Apply to the  Integrated Life Sciences Program . This requires completion of the ILS Background Interest form and a personal statement.

Application deadline

Application packages must be completed by December 4th to be considered for matriculation the following August. Accepted applicants will be notified in March.

Frequently asked questions

How important is gpa.

GPA is important, but we also look at where you went to school and the degree you obtained. Our faculty members must be confident that you will be able to maintain the minimum 3.0 GPA required in graduate level courses.

What should I write in the statement of purpose / personal statement?

This is your chance to “talk” to the Admissions Committee. Express why you want to enroll in our graduate program, what strengths you feel set you apart and highlight your research qualifications.

Who is the Graduate Coordinator? The Graduate Assistant? What’s the difference?

The graduate program assistant is the administrative professional who is responsible for handling incoming applications, maintaining student records, approving forms and submitting information to the Graduate School. You can reach the graduate program assistant at  [email protected] or by phone at 706-542-0389.

Dr. David Peterson is the graduate program coordinator for the department. He chairs the graduate committee that evaluates applications, makes recommendations about the program, and works on programmatic issues such as making application decisions, seeking assistantship funding for new students, and evaluating curriculum changes.

If you need a form signed or a question answered, see the graduate program assistant. If you are seeking a change to the program as a whole, or need assistance troubleshooting some unusual circumstance, see the graduate coordinator.

Who does laboratory rotations and how are they scheduled?

All students applying to our graduate program through the Integrated Life Sciences program at UGA will participate in three different laboratory rotations prior to choosing a major professor. Research rotations provide an opportunity for students to explore the breadth and depth of life science research available at UGA through ILS before formally committing to a thesis project and advisor.

Rotations are scheduled for six weeks in each lab starting in September. After each rotation period, faculty will submit to the ILS graduate coordinator an evaluation form assessing the performance of the student during the rotation.  Students will be able to start in their new labs at the beginning of the spring semester.

Expectations

Students on rotations are expected to spend as much time as possible in the mentoring lab and get immersed into the research culture of the environment. The principal investigator of the lab has the last word about admission of a student into his or her laboratory. In order to make that decision, the professor will assess the student’s performance during the rotation and will have access to the student’s application file.

Are there teaching responsibilities for students on assistantships?

There is no departmental teaching requirement for students who enter our graduate program through the Integrated Life Sciences Program, and whose subsequent stipend support is provided by a grant to their major professor. In the event that a student is supported for a time on a departmental assistantship the student will be required to TA in one of the IDIS courses for each semester they are supported by the department.

Are there teaching opportunities for students who are not on departmental assistantships?

Students who are not required, but wish to have teaching experience, may find teaching opportunities in the Department of Infectious Diseases or in departments with large undergraduate teaching components, such as Microbiology or Cellular Biology.

• Master’s & PhD Programs
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• Comparative Biomedical Sciences
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• PhD in Integrative Physiology & Pharmacology

Contact Information

[email protected]

Department of Infectious Diseases, Department of Population Health, Poultry Diagnostic and Research Center

706-542-5796 office

Department of Infectious Diseases

706-542-0389 office

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Faculty of Medicine

PhD in Virology and Immunology

Graduate 3-503-1-1

Bon à savoir! Ce programme s’adresse autant aux francophones qu’aux anglophones. Consultez la fiche descriptive en français pour en savoir plus.

Contribute to the advancement of knowledge in virology and immunology while conducting innovative, cutting-edge research projects.

About this program

Develop your critical thinking skills and knowledge in research.

Establish yourself as an independent researcher in virology and immunology. In this doctorate program, you will produce an original research project, present your research findings at scientific conferences, and publish them in peer-reviewed journals. You must be able to develop an original research hypothesis and explain the methods used to experimentally validate it. You have the required technical skills for a career in research and excellent critical thinking skills to interpret your results. After completing the program and your subsequent postdoctoral training, you will be positioned to compete for research grants.

• Basic or clinical research program in a wide range of virology and immunology areas
• Practice-oriented training
• Internationally renowned faculty members and research teams
• Collaborative, multidisciplinary, and fertile learning environment

Language accommodations

Although Université de Montréal is a French-language university, many of our research departments are open to creating a bilingual environment for students in graduate-level programs. These departments are aware of the importance of bilingualism in order for graduate students to succeed in their research careers.

Students who are proficient in English are therefore welcome and accepted into graduate-level programs.

The Department of Microbiology, Infectious Diseases and Immunology provides a number of accommodations to make you feel comfortable and help you fulfill the requirements of the program even if French is not your main language of study:

• Access to course notes in both French and English.
• Possibility of taking an equivalent course at an English-language university in Montréal (first session only).
• Choice of giving oral seminar presentations in either French or English.
• Choice of writing exams in French or English.
• Choice of taking the comprehensive exam in either French or English.
• Choice of writing your doctoral thesis or dissertation in French or English.

All laboratories are offered in a bilingual environment. Above all, staff in the program are available to help and support you throughout your studies.

Advantages of studying virology and immunology at UdeM A leading, practice-oriented academic institution

• Develop your potential as a researcher at UdeM, the 4th leading recipient of research grants in Canada
• Discover the largest teaching laboratory in the country with its state-of-the art equipment
• Collaborate with researchers working in various fields of biomedical research

Courses and specifics

For contact information on the program director or student file management technician, or to find out more about the faculty or department hosting the program, please see the page in French .

• Dissertation or thesis track

Reading, critique and discussion of recent scientific articles in the fields of bacteriology, mycology, immunology and virology. Presentation of articles and discussions led by different professors and guest speakers.

Students present their plan, objectives and preliminary results and discuss how they will continue their research project. Question period. Participation in other student seminars.

Students present and interpret their results and discuss how they will continue their research to earn their degree. Question period. Participation in other student seminars.

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More than a century ago, our resiliency has demonstrated the role of microbes in diseases and led to the rational development of vaccines to counter pathogenic microorganisms.

Director, Department of Microbiology, Infectiology and Immunology

Admission requirements

You are eligible only if a faculty member has agreed to supervise your research project or work prior to submitting your admission application.

Be sure to select your choices to display the eligibility conditions that apply to you.

• Official transcripts and two letters of recommendation are required for admission.
• Prospective applicants are strongly encouraged to find a research director prior to submitting their application.

Costs and financial aid

$2,117.74 *

Total for a full-time session of 15 credits

Tuition fees: $1,483.65

Other fees: $634.09

These amounts are estimates and do not account for other expenses, such as costs for insurance, residence, transportation, manuals, etc.

Check your legal status

* These estimates cannot at any time subsitute for an invoice or be used as proof for any reason whatsoever. These calculations are based on the 2024-2025 academic year. Information updated: June 5, 2024

Good news! You may be able to lower this amount!

Under certain conditions, Canadian students who are not residents of Quebec can follow university study programs offered in French while benefiting from the same tuition fees as residents of Quebec.

Check eligibility criteria

$9,753.94 *

Tuition fees: $9,119.85

As an international student, you have access to exemption scholarships granted by UdeM throughout your university program. Note that for ungraduated programs, you must be enrolled as a full-time student for two sessions and reside in Quebec in the case of exclusively online study programs.

Find out about the UdeM exemption scholarship

Available financial assistance

A funding policy ensures a minimum income for graduate students (page in French)

Funding offered for short programs

Funding offered for doctoral programs 

Scholarship program for exemption from additional tuition fees for international students

Scholarships and mobility funding programs offered to students from the Université de Montréal who wish to study abroad

Future prospects Pursuing a career or further studies? The choice is yours!

Your research skills and abilities will secure you a position in academia, industry or a clinical setting.

Need info on the program?

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Tackle the biggest challenges in biology, medicine and health in a world leading research environment, and prepare for your future career.

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PhD/MPhil Medical Virology / Overview

Year of entry: 2025

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We require applicants to hold, or be about to obtain, an Upper Second class Honours degree, or the equivalent qualification gained outside the UK, in a related subject area for entry to a PhD programme. A Lower Second class Honours degree may be considered if applicants also hold a Master's degree with a Merit classification.

Full entry requirements

Apply online

Before applying you must:

• Choose a programme or find a project you want to apply for and check you’re eligible.
• Speak to the listed supervisor about your suitability for their project or programme.
• Understand how your project is funded and, if it is self-funded, consider how you plan on funding it.
• Read our ‘How to apply’ page to find out more and ensure you include all required supporting documents at the time of submission.

Visit our Faculty of Biology, Medicine and Health Postgraduate Research page to find out more.

Programme options

Programme overview

• Undertake research in a field you’re passionate about and join a project addressing leading challenges in the area while working with some of Europe's leading researchers and academics.
• Choose to research at a university ranked and 6th in the UK (QS World University Rankings, 2025) and 2nd in the world for social and environmental impact (THE Impact Rankings, 2024), where 93% of research activity is ‘world leading’ or ‘internationally excellent’ (Research Impact Framework, 2021)
• Access some of the best research facilities in the world at the University, through our industry partners, and at hospitals around Greater Manchester.
• Benefit from dedicated support throughout your PhD journey, from pre-application to graduation and everything in between, through our Doctoral Academy
• Undergo training in transferable skills critical to developing early-stage researchers and professionals through the Doctoral Academy's training programme and progress into a career in research, academia or industry.

Visit our Faculty of Biology, Medicine and Health Postgraduate Research page to find out about upcoming open days and events.

Contact details

Programmes in related subject areas.

Use the links below to view lists of programmes in related subject areas.

• Biosciences

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